101 - Document - Document and Entity Information link:calculationLink link:presentationLink link:definitionLink 103 - Statement - Condensed Consolidated Balance Sheets link:calculationLink link:presentationLink link:definitionLink 104 - Statement - Condensed Consolidated Balance Sheets (Parenthetical) link:calculationLink link:presentationLink link:definitionLink 105 - Statement - Condensed Consolidated Statements of Operations link:calculationLink link:presentationLink link:definitionLink 106 - Statement - Condensed Consolidated Statements of Operations (Parenthetical) link:calculationLink link:presentationLink link:definitionLink 107 - Statement - Condensed Consolidated Statements of Comprehensive Loss link:calculationLink link:presentationLink link:definitionLink 108 - Statement - Condensed Consolidated Statements of Cash Flows link:calculationLink link:presentationLink link:definitionLink 109 - Disclosure - Description of Operations and Summary of Significant Accounting Policies link:calculationLink link:presentationLink link:definitionLink 110 - Disclosure - Collaboration Agreements link:calculationLink link:presentationLink link:definitionLink 111 - Disclosure - Fair Value Measurements link:calculationLink link:presentationLink link:definitionLink 112 - Disclosure - Balance Sheet Components link:calculationLink link:presentationLink link:definitionLink 113 - Disclosure - Stock-Based Compensation link:calculationLink link:presentationLink link:definitionLink 114 - Disclosure - Income Taxes link:calculationLink link:presentationLink link:definitionLink 115 - Disclosure - Related Party Transactions link:calculationLink link:presentationLink link:definitionLink 116 - Disclosure - Description of Operations and Summary of Significant Accounting Policies (Policies) link:calculationLink link:presentationLink link:definitionLink 117 - Disclosure - Description of Operations and Summary of Significant Accounting Policies (Tables) link:calculationLink link:presentationLink link:definitionLink 118 - Disclosure - Collaboration Agreements (Tables) link:calculationLink link:presentationLink link:definitionLink 119 - Disclosure - Fair Value Measurements (Tables) link:calculationLink link:presentationLink link:definitionLink 120 - Disclosure - Balance Sheet Components (Tables) link:calculationLink link:presentationLink link:definitionLink 121 - Disclosure - Stock-Based Compensation (Tables) link:calculationLink link:presentationLink link:definitionLink 122 - Disclosure - Description of Operations and Summary of Significant Accounting Policies - Additional Information (Detail) link:calculationLink link:presentationLink link:definitionLink 123 - Disclosure - Description of Operations and Summary of Significant Accounting Policies - Schedule of Anti-dilutive Securities Excluded from Calculation of Diluted Net Loss per Share (Detail) link:calculationLink link:presentationLink link:definitionLink 124 - Disclosure - Collaboration Agreements - Additional Information (Detail) link:calculationLink link:presentationLink link:definitionLink 125 - Disclosure - Collaboration Agreements - Summary of Revenue Recognized under Agreement (Detail) link:calculationLink link:presentationLink link:definitionLink 126 - Disclosure - Collaboration Agreements - Total Arrangement Consideration Allocated to Deliverables along with Associated Deferred Revenue (Detail) link:calculationLink link:presentationLink link:definitionLink 127 - Disclosure - Fair Value Measurements - Fair Values of Financial Assets Measured on Recurring Basis (Detail) link:calculationLink link:presentationLink link:definitionLink 128 - Disclosure - Fair Value Measurements - Fair Values of Financial Liabilities Carried at Historical Cost (Detail) link:calculationLink link:presentationLink link:definitionLink 129 - Disclosure - Fair Value Measurements - Additional Information (Detail) link:calculationLink link:presentationLink link:definitionLink 130 - Disclosure - Balance Sheet Components - Schedule of Cash and Cash Equivalents (Detail) link:calculationLink link:presentationLink link:definitionLink 131 - Disclosure - Balance Sheet Components - Summary of Amortized Cost, Gross Unrealized Holding Gains or Losses, and Fair Value of Available-for-Sale Investments (Detail) link:calculationLink link:presentationLink link:definitionLink 132 - Disclosure - Balance Sheet Components - Additional Information (Detail) link:calculationLink link:presentationLink link:definitionLink 133 - Disclosure - Balance Sheet Components - Schedule of Accrued Liabilities (Detail) link:calculationLink link:presentationLink link:definitionLink 134 - Disclosure - Stock-Based Compensation - Schedule of Allocated Stock-Based Compensation Expense Related to Options Granted (Detail) link:calculationLink link:presentationLink link:definitionLink 135 - Disclosure - Stock-Based Compensation - Schedule of Weighted-average Assumptions used to Estimate Fair Value of Stock Options and Stock Options Granted (Detail) link:calculationLink link:presentationLink link:definitionLink 136 - Disclosure - Income Taxes - Additional information (Detail) link:calculationLink link:presentationLink link:definitionLink 137 - Disclosure - Related Party Transactions - Additional Information (Detail) link:calculationLink link:presentationLink link:definitionLink